作物学报
  2024,Vol. 50(4): 981-990
  耕作栽培·生理生化
不同播幅对小麦花后叶片光合特性和产量的影响
张振1,赵俊晔2,*,石玉1,张永丽1,于振文1
1山东农业大学农学院 / 小麦育种全国重点实验室 / 农业农村部作物生理生态与耕作重点实验室,山东泰安271018; 2中国农业科学院农业信息研究所 / 农业部农业信息服务技术重点实验室,北京100081
doi: 10.3724/SP.J.1006.2024.31042
Effects of different sowing space on photosynthetic characteristics after anthesis and grain yield of wheat
ZHANG Zhen1,ZHAO Jun-Ye2,*,SHI Yu1,ZHANG Yong-Li1,YU Zhen-Wen1
1College of Agronomy, Shandong Agricultural University / National Key Laboratory of Wheat Breeding / Key Laboratory of Crop Physiology, Ecology and Farming, Ministry of Agriculture and Rural Affairs, Tai’an 271018, Shandong, China; 2 Key Laboratory of Agricultural Information Service Technology, Ministry of Agriculture and Rural Affairs, Agricultural Information Institute, Chinese Academy of Agricultural Sciences, Beijing 100081, China
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摘要: 为了明确不同播幅对小麦籽粒产量的影响及其形成的生理原因,本研究于2019—2020年和2020—2021年冬小麦生长季,在山东省济宁市兖州区小孟镇史家王子村小麦试验站大田试验条件下设置2种播幅处理:处理1是播幅为8 cm (B1);处理2是播幅为3 cm (B2)。研究了不同播幅对小麦光合特性、冠层光截获特性、干物质积累与转运和籽粒产量的影响。试验结果表明:B1处理开花后叶面积指数和冠层光合有效辐射截获率显著高于B2处理,其冠层光合有效辐射透射率显著低于B2处理;B1处理开花后旗叶叶绿素相对含量、净光合速率、蒸腾速率和气孔导度均显著高于B2处理,其胞间二氧化碳浓度显著低于B2处理;B1处理开花期和成熟期干物质积累量、开花后干物质在籽粒中的分配量、成熟期籽粒干物质积累量均显著高于B2处理;B1处理穗粒数、千粒重均显著高于B2处理;与B2处理相比,B1处理的两年平均籽粒产量和光能利用率分别高6.12%和7.71%。综上所述,播幅为8 cm的B1处理通过塑造了合理的冠层结构,改善了开花后叶片的光合性能,有利于开花后植株的光合物质生产,从而获得了最高的籽粒产量和光能利用率,为本试验条件下的最优处理。研究为小麦宽幅播种节水高产高效栽培技术提供了理论依据。
关键词: 小麦,  光合特性,  干物质积累与转运,  籽粒产量 
Abstract:

With the development of industrialization process in society, the problem of cadmium (Cd) pollution in soil is increasing. However, Nicotiana tabacum has a strong Cd enrichment capacity in leaves, which seriously affects its economic value. To investigate the mechanism by which tobacco responds to Cd stress, tobacco leaves were harvested from the culture solution with Cd concentrations of 0 μmol L-1 and 500 μmol L-1 for subsequent transcriptome sequencing. In this study, a total of 76.94 Gb clean data was obtained, with Q30 base percentage exceeding 95.43%. The results showed that 7735 differentially expressed genes (DEGs) were screened under Cd stress conditions, including 4833 up-regulated genes and 2902 down-regulated genes. The reliability of transcriptome data was verified by qRT-PCR analysis to detect the expression patterns of candidate gene. Gene ontology (GO) annotation as well as Kyoto encyclopedia of genes and genomes (KEGG) pathway analysis were performed on differentially expressed transcripts. GO functional enrichment revealed that the differentially expressed genes were mainly distributed in metabolic processes, response to stimulus, cellular anatomical entity, catalytic activity, and transcription regulator activity. Meanwhile, KEGG analysis showed that the up-regulated differentially expressed genes were mainly involved in biosynthesis of amino acids, carbon metabolism, oxidative phosphorylation, and citrate cycle. Down-regulated differentially expressed genes were primarily enriched in photosynthesis, biosynthesis of secondary metabolites, metabolic pathways, and plant hormone signal transduction. Further analysis of plant hormone signal transduction pathways revealed that there were eight plant hormone pathways involved in response to cadmium stress in tobacco, and the relative expression patterns of different hormone gene member were also different. Experimental results from plant hormone application on tobacco leaves demonstrated that the regulation of gibberellins, brassinosteroids, and jasmonic acid pathways played roles in tobacco's response to cadmium stress. The experimental results of Arabidopsis hormone signal mutant showed that plants respond to cadmium stress by regulating ethylene, gibberellin, brassinosteroid, and jasmonic acid pathways. In conclusion, this study not only explores the regulatory network of tobacco resistance to Cd stress, but also lays a theoretical foundation for the genetic improvement of crop resistance.

Key words: wheat,  photosynthetic characteristics,  dry matter accumulation and transport,  grain yield 
收稿日期: 2023-02-16
基金资助: 本研究由国家自然科学基金项目(32172114, 31601243), 财政部和农业农村部国家现代农业产业技术体系建设专项(CARS-03)和泰山学者工程专项经费资助。